| نقل قول | Ward, M., 1941 New Brachyura from the Gulf of Davao, Mindanao, Philippine Islands. American Museum Novitates (1104):1-15. |
| مرجع | 2897 Ward, M., 1941 |
| Page | 10 |
| Named Used as Valid | Leptodius sanguineus philippinensis |
| Locality | Davao Gulf |
| Distribution | Philippines. |