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References used for species in the Family Onuphidae
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[ n = 118 ] 
Al-Yamani, F.Y., V. Skryabin, N. Boltachova, N. Revkov, M. Makarov, V. Grinstov and E. Kolesnikova, 2012 Illustrated Atlas on the Zoobenthos of Kuwait. Kuwait Institute for Scientific Research.
Arana, I.L. and O.D. Díaz, 2006 Polychaeta (Annelida) associated with Thalassia testudinum in the northeastern coastal waters of Venezuela. Revista de Biología Tropical 54(3):971-978.
Augener, H., 1924 Polychaeta von Neuseeland. I. Errantia. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening I Kjøbenhavn 75:241-441, 11 figs.
Australian Biological Resources Study, 2000 Australian Faunal Directory. World Wide Web electronic publication.
Bergmann, M., N. Langwald, J. Ontrup, T. Soltwedel, I. Schewe, M. Klages and T.W. Nattkemper, 2011 Megafaunal assemblages from two shelf stations west of Svalbard. Marine Biology Research 7:523-539.
Bisby, F.A., M.A. Ruggiero, K.L. Wilson, M. Cachuela-Palacio, S.W. Kimani, Y.R. Roskov, A. Soulier-Perkins and J. van Hertum, 2005 Species 2000 & ITIS Catalogue of Life: 2005 Annual Checklist. CD-ROM; Species 2000: Reading, U.K.
Bisby, F.A., Y.R. Roskov, T.M. Orrell, D. Nicolson, L.E. Paglinawan, N. Bailly, P.M. Kirk, T. Bourgoin and J. van Hertum (eds.), 2008 Species 2000 & ITIS Catalogue of Life: 2008 Annual Checklist. CD-ROM; Species 2000: Reading, U.K.
Boletzky, S.V. and R.T. Hanlon, 1983 A review of the laboratory maintenance, rearing and culture of cephalopod molluscs. 44:147-187. In C.F.E. Roper, C.C. Lu and F.G. Hochberg (eds.) Memoirs of the National Museum of Victoria: Proceedings of the Workshop on the Biology and Resource Potential of Cephalopods, Melbourne, Australia, 9-13 March, 1981.
Bonsdorff, E. and T.H. Pearson, 1999 Variation in the sublittoral macrozoobenthos of the Baltic Sea along environmental gradients: A functional-group approach. Australian Journal of Ecology 24:312-326.
Branch, G.M., C.L. Griffiths, M.L. Branch and L. Beckley, 2005 Two oceans: a guide to marine life in southern Africa. Struik Publishers, Cape Town, 360 p.
Callaway, R., J. Alsvag, I. de Boois, J. Cotter, A. Ford, H. Hinz, S. Jennings, I. Kroncke, J. Lancaster, G. Piet, P. Prince and S. Ehrich, 2002 Diversity and community structure of epibenthic invertebrates and fish in the North Sea. ICES Journal of Marine Science 59:1199-1214.
Carlier, A., P. Riera, J.-M. Amouroux, J.-Y. Bodiou and A. Grémare, 2007 Benthic trophic network in the Bay of Banyuls-sur-Mer (northwest Mediterranean, France): an assessment based on stable carbon and nitrogen isotopes analysis. Estuarine, Coastal and Shelf Science 72(1-2):1-15.
Chao, M., Y. Shi, W. Quan, X. Shen, C. An, Q. Yuan and H. Huang, 2012 Distribution of benthic macroinvertebrates in relation to environmental variables across the Yangtze River Estuary, China. Journal of Coastal Research 28(5):1008-1019.
Chava, A.I., A.A. Udalov, A.A. Vedenin, M.I. Simakov, S.A. Shchuka and V.O. Mokievsky, 2017 Benthic fauna of Tsivolki Bay (Novaya Zemlya Archipelago, Kara Sea). Oceanology 57(1):144-153.
Costello, D.P. and C. Henley, 1971 Methods for obtaining and handling marine eggs and embryos. Marine Biological Laboratory, Woods Hole, MA (Second Edition).
Cusson, M., P. Archambault and A. Aitken, 2007 Biodiversity of benthic assemblages on the Arctic continental shelf: historical data form Canada. Mar. Ecol. Prog. Ser. 331:291-304.
Dauvin, J.C. and M. Joncourt, 1989 Energy values of marine benthic invertebrates from the western English Channel. Journal of the Marine Biological Association of the U.K. 69:589-595.
Dean, H.K., 2009 Polychaetes and echiurans. p. 181-192 Wehrtmann, I.S. and Cortés, J. 2009. Marine biodiversity of Costa Rica, Central America. Springer 538pp.
Delle Chiaje, S., 1841 I. Anellosi Amfitritici. II. Anellosi Talassemici. III. Anellosi Lombrici. IV. Anellosi Nereidici. Descrizione e notomia degli animali invertebrati della Sicilia citeriore osservati vivi negli anni 1822-1830 3:69-104.
Drumm, D.T., K.P. Maslenikov, R. Van Syoc, J.W. Orr, R.R. Lauth, D.E. Stevenson and T.W. Pietsch, 2016 An annotated checklist of the marine macroinvertebrates of Alaska. NOAA Professional Paper NMFS 19, 289 p. doi:10.7755/PP.19
Ehlers, E., 1908 Die bodensässigen Anneliden aus den Sammlungen der deutschen Tiefsee-Expeditionen. In Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer Valdivia 1898-1899. Chun, C. (ed.):16 (1): 1-167 + 1-23 plates. Gustav Fischer Jena.
Ellis, J.R., G. Burt and S.I. Rogers, 2007 Epifaunal sampling in the Celtic Sea. ICES ASC 2007 Theme Session A23.
FAO-FIES, 2007 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from, [accessed 03/07/2008].
Fauchald, K., 1982 Revision of Onuphis, Nothria, and Paradiopatra (Polychaeta: Onuphidae) based upon type material. Smithsonian Institution Press.
Fauchald, K. and P.A. Jumars, 1979 The diet of worms: a study of polychaete feeding guilds. Oceanography and Marine Biology 17:193-284.
Fauchald, K., A. Granados-Barba and V. Solis-Weiss, 2009 Polychaeta (Annelida) of the Gulf of Mexico. pp. 711-744 In Felder, D.L.; Camp, D.K. (eds.) Gulf of Mexico - Origins, Waters, and Biota. Biodiversity. Texas A&M University Press, College Station, Texas
Fauvel, P., 1932 Annelida Polychaeta of the Indian Museum, Calcutta. Memoirs of the Indian Museum 12(1):1-262.
Fox, R.S. and E.E. Ruppert, 1985 Shallow-water marine benthic invertebrates of South Carolina: Species identificattion, community composition and symbiotic association. University of South Carolina Press, Columbia, SC (USA).
Franco-Gordo, M., 2013 Inventario de biodiversidad de la costa sur de Jalisco y Colima. Universidad de Guadalajara.
Fuhrmann, M.M., T. Pedersen and E.M. Nilssen, 2017 Trophic niche of the invasive red king crab Paralithodes camtschaticus in a benthic food web. Marine Ecology Progress Series 565:113-129.
Gallardo, V.A., 1968 Polychaeta from the Bay of Nha Trang, South Viet Nam. Scripps Instn. Oceanogr., Univ. Calif. NAGA Exped. Rep. 4(3):35-279.
Gambi, M.C. and S. Mariani, 1999 Polychaetes of the soft bottoms of the Straits of Magellan collected during the Italian oceanographic cruise in February-March 1991. Scientia Marina 63(Suppl 1):233-242.
Garilao, N., 2014 Baltic Sea species list. Personal communication, 25 April 2014. Provided excel file.
Gaudêncio, M.J. and H.N. Cabral, 2007 Trophic structure of macrobenthos in the Tagus estuary and adjacent coastal shelf. Hydrobiologia 587(1):241-251.
Gibbs, P.E., 1978 Macrofauna of the intertidal sand flats on low wooded islands, Northern Great Barrier Reef. Philosophical Transactions of the Royal Society of London, Series B 284(999):81-97.
Gotshall, D.W., 2005 Guide to marine invertebrates: Alaska to Baja California, 2nd ed. (revised). Sea Challengers. 117 p.
Gusso, C.C., M.F. Gravina and F.R. Maggiore, 2001 Temporal variations in soft bottom benthic communities in central Tyrrhenian Sea (Italy). Archo Oceanogr. Limnol. 22:175-182.
Hartman, O., 1974 Polychaetous annelids of the Indian Ocean including an account of species collected by members of the International Indien Ocean Expeditions, 1963-64 and a catalogue of bibliography of the species from India. Journal of the Marine Biological Association of India 16(1):191-252 + 13 figures.
Hernández-Alcátara, P. and V. Solís-Weiss, 1991 New records of errantiate polychaetous annelids from the Gulf of California. Bulletin of Marine Science 48(2):251-260.
Hogg, O.T., D.K.A. Barnes and H.J. Griffiths, 2011 Highly diverse, poorly studied and uniquely threatened by climate change: an assessment of marine biodiversity on South Georgia's continental shelf. PLoS ONE 6(5):e19795.
Hong, S.Y., K.Y. Park, C.W. Park, C.H. Han, H.L. Suh, S.G. Yun, C.B. Song, S.G. Jo, H.S. Lim, Y.S. Kang, D.J. Kim, C.W. Ma, M.H. Son, H.K. Cha, K.B. Kim, S.D. Choi, K.Y. Park, C.W. Oh, D.N. Kim, H.S. Shon, J.N. Kim et al., 2006 Marine invertebrates in Korean coasts. Republic of Korea, Academy Publishing Company, Inc. 479 p.
Hsieh, H.L. and K.H. Chang, 1991 Habitat characteristics and occurrence of the spionid Pseudopolydora sp. on the tube-caps of the onuphid Diopatra bilobata (Polychaeta: Spionidae, Onuphidae). Bull. Inst. Zool. Acad. Sin. (Taipei) 30(4):331-339.
Jia, X., Z. Li, C. Li, Y. Qiu and J. Gan, 2004 Studies on the ecosystem and commercial fisheries resources in commercial zone and the continental shelf in South China Sea. Science Publishing, Beijing, 647 p.
Kedra, M., S. Gromisz, R. Jaskula, J. Legezynska, B. Maciejewska, E. Malec, A. Opanowski, K. Ostrowska, M. Weodarska-Kowalczuk and J. Weslawski, 2010 Soft bottom macrofauna on All Taxa Biodiversity Site: Hornsund (77°N, Svalbard). Polish Polar Research 31(4):309-326.
Kirkegaard, J.B., 1995 Bathyal and abyssal polychaetes (errant species). Scientific Results of the Danish Deep-Sea Expedition Round the World 1950-52. Galathea 2 Reports 17:7-56.
Lamb, A., D. Gibbs and C. Gibbs, 2011 Strait of Georgia biodiversity in relation to bull kelp abundance. Pacific Fisheries Resource Conservation Council, 111p.
Lechapt, J.P., 1997 Two new species of Hyalinoecia (Polychaeta, Onuphidae) from deep zones off New Caledonia (Southwest Pacific Ocean). Bulletin of Marine Science 60(2):306-312.
López-Jamar, E., G. González and J. Mejuto, 1986 Temporal changes of community structure and biomass in two subtidal macroinfaunal assemblages in La Coruña bay, NW Spain. Hydrobiologia 142:137-150.
M'Intosh, W.C., 1885 Report on the Annelida Polychaeta dredged H.M.S. Challenger during the years 1873-76. In Report of the scientific results of the voyage of H.M.S. Challenger during the years 1873-76: a summary of the scientific results, Zoology 12:1-590.
Macdonald, T.A., B.J. Burd, V.I. Macdonald and A. van Roodselaar, 2010 Taxonomic and feeding guild classification for the marine benthic macroinvertebrates of the Strait of Georgia, British Columbia. Canadian Technical Report of Fisheries and Aquatic Sciences 2874:62p.
MarBEF, 2006 MarBEF Data System.[accessed June 26, 2006], 2050
Martin, D. and J. Gil, 2010 Checklist of class Polychaeta (Phylum Annelida). pp. 199-236 In Coll, M., et al., 2010. The biodiversity of the Mediterranean Sea: estimates, patterns, and threats. PLoS ONE 5(8):36pp.
McDaniel, N. and D. Swanston, 2013 Observations on the Gorgonian Coral Primnoa pacifica at the Knight Inlet sill, British Columbia.,-may-1-13.pdf [Accessed 05/01/2017].
Miloslavich, P., J.M. Díaz, E. Klein, J.J. Alvarado, C. Díaz, J. Gobin, E. Escobar-Briones, Motta, J.J. Cruz, E. Weil, J. Cortés, A.C. Bastidas, R. Roberston, F. Zapata, A. Martín, J. Castillo, A. Kazandjian and M. Ortiz, 2010 Marine biodiversity in the Caribbean: regional estimates and distribution patterns. PLoS ONE 5(8):e11916.
Monro, C.C.A., 1937 The John Murray Expedition 1933-34. Polychaeta. Scientific Reports, Zoology 4(8):243-321 + 28 figures.
Moreno, R.A., R.D. Sepúlveda, E.I. Badano, S. Thatje, N. Rozbaczylo and F.D. Carrasco, 2008 Subtidal macrozoobenthos communities from northern Chile during and post El Niño 1997-1998. Helgol Mar Res 62 (Suppl 1):S45-S55.
Murray, J., 1896 On the deep and shallow-water marine fauna of the Kerguelen region of the Great Southern Ocean. Challenger Expedition 343-500 p.
Ng, P.K. and N. Sivasothi, 1999 Guide to the Mangroves of Singapore II (Animal Diversity). Singapore Science Centre, pp. 1-158.
OBIS, 2016 Diopatra neapolitana Delle Chiaje, 1841. OBIS Search Interface. [Accessed 4/29/2016].
OBIS, 2016 Kinbergonuphis investigatoris (Fauvel, 1932). OBIS Search Interface. [Accessed 4/29/2016].
OBIS, 2016 Onuphis eremita Audouin & Milne Edwards, 1833. OBIS Search Interface. [Accessed 4/8/2016].
Paxton, H. and J.H. Bailey-Brock, 1986 Diopatra dexiognatha, a new species of Onuphidae (Polychaeta) from Oahu, Hawaiian Islands. Pacific Science 40:1-4.
Paxton, H. and L.M. Chou, 2000 Polychaetous annelids from the South China Sea. The Raffles Bulletin of Zoology 8:209-232.
Piepenburg, D., N.V. Chernova, C.F. von Dorrien, J. Gutt, A.V. Neyelov, E. Rachor, L. Saldanha and M.K. Schmid, 1996 Megabenthic communities in the waters around Svalbard. Polar Biol. 16:431-446.
Pires, A., F. Gentil, V. Quintino and A.M. Rodrigues, 2012 Reproductive biology of Diopatra neapolitana (Annelida, Onuphidae), an exploited natural resource in Ria de Aveiro (Northwestern Portugal). Marine Ecology 33(1):56-65.
Plotkin, P.T., M.K. Wicksten and A.F. Amos, 1993 Feeding ecology of the loggerhead sea turtle Caretta caretta in the northwestern Gulf of Mexico. Mar. Biol. 115:1-5.
Robinson, L.A., S.P.R. Greenstreet, H. Reiss, R. Callaway, J. Ccraeymeersch, I. de Boois, S. Degraer, S. Ehrich, H.M. Fraser, A. Goffin, I. Kröncke, L. Lindal Jorgenson, M.R. Robertson and J. Lancaster, 2010 Length-weight relationships of 216 North Sea benthic invertebrates and fish. J. Mar. Biol. Ass. U.K. 90(1):95-104.
Ruppert, E.E., R.S. Fox and R.D. Barnes, 2004 Invertebrate Zoology. A functional evolutionary approach. 7th Ed. Brooks/Cole, Thomson Learning learning, Inc. 990 p.
Salazar-Vallejo, S.I., 1996 Lista de species y bibliografía de Poliquetos (Polychaeta) del Gran Caribe. Anales Inst. Biol. Univ. nac. Autón, México, Ser. Zool. 67(1):11-50.
Salazar-Vallejo, S.I. and M.H. Londoño-Mesa, 2004 Lista de especies y bibliografía de poliquetos (Polychaeta) del Pacífico Oriental Tropical. Anales del Instituto de Biología, Universidad Nacional Autónoma de México, Serie Zoología 75(1):9-97.
Schoch, G.C. and M. Dethier, 2001 The intertidal biota of Puget Sound gravel beaches. Report for the Washington State Department of Natural Resources Nearshore Habitat Program, 48p.
Shen, S., 1987 A new species of Onuphis (Polychaeta: Onuphidae). Symposium on Research Reports on the Zanomu Ansha of Nansha Islands of China, Science Press, Beijing:221-226.
Shin, P.K.S., 1982 Some polychaetous annelids from Hong Kong waters. In B.S. Morton and C.K. Tseng (eds.), Proceedings of the first International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong 1980:161-172. Hong Kong University Press, Hong Kong.
Shin, P.K.S. and G.B. Thompson, 1982 Spatial distribution of the infaunal benthos of Hong Kong. Mar. Ecol. Prog. Ser. 10:37-47.
Sicinski, J. and P. Gillet, 2002 Biogeographical affinities of polychaetes from Îles Crozet. Antarc. Sci. 14(4):353-363.
Sirenko, B., S. Denisenko, H. Deubel and E. Rachor, 2004 Deep water communities of the Laptev Sea and adjacent parts of the Arctic Ocean. Russian Academy of Sciences, Zoological Institute Exploration of the Fauna of the Seas 54(62):28-73.
Smithsonian Institution, 2016 Americonuphis reesei Fauchald, 1973. [Accessed 07/07/2016].
Smithsonian Institution, 2016 Diopatra neotridens Kinberg. [Accessed 07/07/2016].
Smithsonian Institution, 2016 Diopatra papillata Fauchald. [Accessed 07/07/2016].
Smithsonian Institution, 2016 Diopatra splendidissima Kinberg. [Accessed 07/07/2016].
Smithsonian Institution, 2016 Diopatra tridentata Hartman, 1944. [Accessed 07/07/2016].
Smithsonian Institution, 2016 Hyalinoecia juvenalis Moore. [Accessed 07/07/2016].
Smithsonian Institution, 2016 Kinbergonuphis difficilis (Fauchald, 1982). [Accessed 07/14/2016].
Smithsonian Institution, 2016 Kinbergonuphis orensanzi (Fauchald, 1982). [Accessed 07/14/2016].
Smithsonian Institution, 2016 Kinbergonuphis simoni (Santos et al., 1981). [Accessed 07/07/2016].
Smithsonian Institution, 2016 Longibrachium atlanticum (Day). [Accessed 07/14/2016].
Smithsonian Institution, 2016 Mooreonuphis dangrigae (Fauchald, 1980). [Accessed 07/14/2016].
Smithsonian Institution, 2016 Mooreonuphis jonesi Fauchald. [Accessed 07/14/2016].
Smithsonian Institution, 2016 Mooreonuphis nebulosa (Moore, 1911). [Accessed 07/07/2016].
Smithsonian Institution, 2016 Mooreonuphis pallidula (Hartman, 1965). [Accessed 07/07/2016].
Smithsonian Institution, 2016 Nothria conchylega (Sars, 1835). [Accessed 07/07/2016].
Smithsonian Institution, 2016 Onuphis eremita Hartman, 1951. [Accessed 05/23/2016].
Smithsonian Institution, 2016 Onuphis pulchra Fauchald. [Accessed 07/14/2016].
Smithsonian Institution, 2016 Onuphis texana Fauchald. [Accessed 07/14/2016].
Smithsonian Institution, 2016 Paradiopatra fragosa (Ehlers, 1887). [Accessed 07/07/2016].
Smithsonian Institution, 2016 Paradiopatra hartmanae (Kirkegaard, 1980). [Accessed 07/14/2016].
Smithsonian Institution, 2016 Paradiopatra pycnobranchiata (McIntosh, 1885). [Accessed 07/14/2016].
Solís-Weiss, V., Y. Bertrand, M. Helléouet and F. Pleijel, 2004 Types of polychaetous annelids at the Muséum national d'Histoire naturelle, Paris. Zoosystema 26(3):377-384.
Southern Ocean Mollusc Database (SOMBASE), 2011 British Antarctic Survey, SOMBASE. Accessed through GBIF data portal,, 2011-07-15.
Steimle, F.W. Jr. and R.J. Terranova, 1988 Energy contents of Northwest Atlantic continental slope organisms. Deep Sea Res. 35:415-423.
Tan, L.T. and L.M. Chou, 1996 Diopatra bulohensis, a new species of Onuphidae (Polychaeta) from Sungei, Buloh, Singapore. Raffles Bull. Zool. 44(2):357-362.
Tan, L.T. and L.M. Chou, 1993 Checklist of polychaete species from Singapore waters (Annelida). Raffles Bulletin of Zoology 41(2):279-295.
Tan, L.T. and L.M. Chou, 2005 New eunicid polychaete of the genus Nauphanta Kinberg 1865 (Annelida: Eunicidae) from Singapore. Zoological Science.
Thayer, G.W., W.E. Schaaf, J.W. Angelovic and M.W. LaCroix, 1973 Caloric measurements of some estuarine organisms. Fish. Bull. 71:289-296.
The Danish Biodiversity Information Facility, 2007 Galathea II, Danish Deep Sea Expedition 1950-52. (accessed through GBIF data portal, 13/09/2007).
Tracey, D.M., O.F. Anderson, M.R. Clark and M.D. Olver (Comps.), 2005 A guide to common deepsea invertebrates in New Zealand waters. New Zealand Aquatic Environment and Biodiversity Report No. 1. 160 p.
Treadwell, A.L., 1939 Four new species of polychaetous annelids collected by the United States fisheries steamer "Albatross" during the Philippine expedition of 1907-1910: contribution to the biology of the Philippine Archipelago and adjacent regions. p. 313-321. In U.S. National Museum. Papers on Philippine diatoms, annelids, hydroids, echinoids, and mollusks: contribution to the biology of the Philippine Archipelago and adjacent regions. Smithsonian Institution United States National Museum Bulletin 100(6):567 p.
Vittor, B.A., 2002 Gray's Reef benthic macroinvertebrate community assessment, April 2001. Final Report to NOAA under contract No. 50-DGNC-0-90024, NOAA NOS, Charleston, SC.
Vittor, B.A., 2001 Florida Bay macroinvertebrate community assessment, August 2000. NOAA, NOS, Charleston, SC 29412.
Wacasey, J.W., 1975 Biological productivity of the Southern Beaufort Sea: zoobenthic studies. Beaufort Sea Technical Report 12b, 42p.
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Weslawski, J.M., M. Wlodarska-Kowalczuk and J. Legezynska, 2003 Occurrence of soft bottom macrofauna along the depth gradient in High Arctic, 79° N. Polish Polar Research 24(1):73-88.
Wu, B., R. Sun and M. Chen, 1980 Zoogeographical studies on Polychaeta from the Xisha Islands and its adjacent waters. Acta Oceanol. Sin. 2(1):111-130.
Wu, Q., 1993 Polychaete ecology in soft-bottom in western Taiwan Strait. Journal of Oceanography in Taiwan Strait 12(4):324-334.
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