| شماره مرجع | Márquez, M.R., 1990 |
| بوم سازگان _ اكوسيستم | Nicoya |
| وضعیت | native |
| فراواني | |
| Lifestage | adults |
| Regional database | |
| Remarks | Known from Tortuguero as one of the nesting grounds (Ref. 1397); habitat for major nesting colonies (Ref. 66074). Nesting season occurs from April to October with a peak between June and September (Ref. 1397). C: Ref. 83945; M: Ref. 1397; O: Refs. 1397, |